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Alex, Marty, Gloria and Melman are still trying to get back to the Big Apple and their beloved Central Park zoo, but first they need to find the penguins. They travel to Monte Carlo where they attract the attention of Animal Control after gate crashing a party and are joined by the penguins, King Julian and Co., the monkeys and a new arrival: a performing llama. How do a lion, zebra, hippo, giraffe, four penguins, two monkeys, three lemurs and a llama travel through Europe without attracting attention? They join a traveling circus.